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East Midlands Railway is a train operating company that provides passenger services across the East Midlands region of England. It was formed in 2019, taking over the franchise previously operated by East Midlands Trains. The company operates services on routes including London St Pancras to Sheffield, Nottingham to Liverpool, and Norwich to Liverpool. East Midlands Railway also operates local services within the East Midlands, connecting towns and cities such as Derby, Leicester, and Lincoln.

Greater Anglia, which is operated by the Transport UK Group, provides passenger rail services in the East of England. It operates services on the Great Eastern Main Line from London Liverpool Street to destinations such as Norwich, Ipswich, and Colchester, as well as on the West Anglia Main Line from London Liverpool Street to Cambridge and Stansted Airport. Greater Anglia also operates regional services in Norfolk, Suffolk, Essex, and Cambridgeshire. The Department for Transport has awarded the operating contract for services until 2025 to Abellio East Anglia Limited.

London North Eastern Railway (LNER) is a train operating company in the United Kingdom that operates intercity services on the "East Coast Main Line" between London, Yorkshire, the North East and Scotland. It was formed in 2018 as a result of the UK government's decision to terminate the franchise of Virgin Trains East Coast. LNER is owned by the Department for Transport and managed by the operator of the East Coast Partnership, a joint venture between infrastructure company Network Rail and train operator FirstGroup. LNER operates a fleet of electric and diesel trains, including the Azuma, which is the fastest train in the UK.

London Overground is a suburban rail network in London, England. It was created in 2007 by the merger of the former Silverlink Metro service with other rail services. The network consists of 112 stations and serves 23 London boroughs. London Overground operates on these routes, which include Highbury & Islington to West Croydon/Clapham Junction/Crystal Palace, Richmond/Clapham Junction to Stratford, Gospel Oak to Barking Riverside, Watford Junction to Euston, Liverpool Street to Enfield Town/Cheshunt, Liverpool Street to Chingford, and Romford to Upminster. Overground is operated by Arriva Rail London under a concession from Transport for London.

The London Underground, also known as "the Tube", is a rapid transit system that serves London and some parts of the surrounding counties. It is the oldest underground railway network in the world, with the first line opening in 1863. The Tube is a vital part of London's transportation infrastructure, carrying over five million passengers per day. It is operated by Transport for London (TfL) and is known for its iconic map, distinctive roundel logo, and deep-level tunnels.

Northern is a train operating company that provides passenger services across the north of England. Northern is owned by Northern Trains Ltd, a division of "DOHL," the public sector holding company for the Department of Transportation. It is owned by Arriva UK Trains and operates over 2,500 services every weekday, serving cities such as Manchester, Liverpool, Leeds, and Newcastle.

Thameslink is a railway service in the United Kingdom that runs through central London, connecting Bedford in the north and Brighton in the south. It is operated by Govia Thameslink Railway and serves over 200 stations, including major transport hubs such as St Pancras International, London Bridge, and Gatwick Airport. Thameslink trains run every few minutes during peak hours and offer a range of services, from fast express trains to stopping services that serve local communities. The service has undergone significant upgrades in recent years, including the introduction of new trains and the completion of major infrastructure projects such as the Thameslink Programme.

TransPennine Express is a train operating company in the United Kingdom that provides intercity train services across the North of England and Scotland. The company operates services between major cities such as Manchester, Liverpool, Leeds, Newcastle, Edinburgh, and Glasgow. Transpennine Express offers a range of ticket options, including advance purchase tickets, out-of-season tickets, and flexible tickets. The company also provides onboard facilities such as free Wi-Fi, power sockets, and refreshments.

West Midlands Trains is a train operating company. The company is operated by a joint venture between Abellio, JR East, and Mitsui and runs under a franchise agreement with the Department for Transport. West Midlands Trains operates a fleet of over 400 trains, serving over 150 stations across the region, including Birmingham, Coventry, Wolverhampton, and Stoke-on-Trent. The firm provides a variety of services, including long-distance trips to places like London and Liverpool as well as local and regional trains.

National Rail is a network of railway companies that operate in England, Scotland, and Wales. It encompasses a network of train operators, including Train Operating Companies, open access operators, freight operating companies, and Network Rail. National Rail is managed and supported by the Rail Delivery Group (RDG), which provides essential business services to ensure the smooth operation and coordination of the railway industry.